The 5 possible Exchange 2007 server roles are:

Unified Messaging
Edge Transport
Client Access
Mailbox
Hub Transport

For an indepth look at these roles, click one of the Exchange 2007 roles above.

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The Exchange 2007 unified messaging role:

Merges Voice over IP with exchange 2007
Allows users to check voice mail through OWA (Outlook Web Access

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Exchange 2007 client access server is:

Provides OWA (Outlook Web Access)
Outlook mobile access

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The Exchange 2007 edge transport role contains:

Anti spam protection
Anti virus protection
When used email hits the edge transport before going to the hub transport server
Can not exist on a server with any other server role.  (IE: a Hub Transport and Edge Transport server can not exist on the same physical machine)
Installed on the edge of your [...]

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The Exchange 2007 Hub Transport role is:

Responsible for internal mail flow
Installed in Active Directory
Varies if you are using an edge transport server as well.

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The Exchange 2007 mailbox role provides and uses the following:

Holds Exchange databases
Hold mailboxes and public folders
Part of an Active Directory Domain
Requires plenty of space in Active Directory
Allows a maximum of 50 mail storage groups (in Exchange 2007 enterprise)
Allows maximum of 50 mail databases across all storage groups (in Exchange 2007 enterprise)

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