The Exchange 2007 mailbox role provides and uses the following:

Holds Exchange databases
Hold mailboxes and public folders
Part of an Active Directory Domain
Requires plenty of space in Active Directory
Allows a maximum of 50 mail storage groups (in Exchange 2007 enterprise)
Allows maximum of 50 mail databases across all storage groups (in Exchange 2007 enterprise)

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If you migrated from Exchange 2003 to Exchange 2007 and still have some residual mailboxes left behind on the old exchange 2003 server, this Power Shell cmdlet will be very useful to you.  It will list all users the old exchange 2003 server and their respective mailstores.
Get-Mailbox -OrganizationalUnit OUNAME -Server OLDSERVER | Format-List Alias, ServerLegacyDN
Where [...]

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